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What is the difference between these calls?

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None. The Assembly.CreateInstance actually calls Activator.CreateInstance under the hood.

Using Reflector on Assembly.CreateInstance:

public object CreateInstance(string typeName, bool ignoreCase, BindingFlags bindingAttr, Binder binder, object[] args, CultureInfo culture, object[] activationAttributes)
{
    Type type = this.GetType(typeName, false, ignoreCase);
    if (type == null)
    {
        return null;
    }
    return Activator.CreateInstance(type, bindingAttr, binder, args, culture, activationAttributes);
}

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